Briefly mention the contribution of $T.H.$ Morgan in genetics.

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(N/A) $T.H.$ Morgan's work was primarily based on the fruit fly, $Drosophila$ $melanogaster$.
He formulated the chromosomal theory of linkage.
He defined linkage as the physical co-existence of two or more genes on the same chromosome.
He performed dihybrid crosses in $Drosophila$ to demonstrate that linked genes are inherited together and are located on the $X$-chromosome.
His experiments also proved that tightly linked genes show very low recombination frequencies, whereas loosely linked genes show higher recombination frequencies.

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Morgan hybridized yellow-bodied,white-eyed females to brown-bodied,red-eyed males and intercrossed their $F_{1}$ progeny. He observed that:
$(a)$ $F_{2}$ ratio deviated significantly from the $9 : 3 : 3 : 1$ ratio.
$(b)$ Both genes did not segregate independently of each other.
$(c)$ Recombinant types are not obtained in $F_{2}$ generation.
$(d)$ Both genes segregate independently of each other.
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