Among bryophytes, lichens and fern which one is a pioneer species in a xeric succession ?

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The species that invade a bare area are called pioneer species which later pave way to some small plants like bryophytes which are with time succeeded by bigger plants.

Lichen produces Lichen acid and carbonic acid and that weathers surface of rock and releases required minerals for growth. In weathered rock, soil particles are collected by wind and provide surface for bryophytes and pterido phytes (Fern Nephrolepis).

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