If the population density at time $t$ is $N_t$,then the population density at time $t+1$ is given by:
- A$N_t = N_{t+1} + [(B + I) - (D + E)]$
- B$N_t = N_{t+1} + [(B + I) - (D - E)]$
- C$N_{t+1} = N_t + [(B + I) - (D + E)]$
- D$N_{t+1} = N_t + [(B + I) - (D - E)]$
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