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| Column $I$ | Column $II$ |
| $1$. August Weismann | $a$. Theory of origin of species |
| $2$. Hugo de Vries | $b$. Theory of chemical evolution |
| $3$. Charles Darwin | $c$. Theory of continuity of germplasm |
| $4$. Oparin and Haldane | $d$. Mutation theory |
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