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| स्तंभ-$I$ | स्तंभ-$II$ |
| $P. +,+$ | $I. \text{सहपरोपकारिता}$ |
| $Q. -,-$ | $II. \text{परजीविता}$ |
| $R. -,0$ | $III. \text{स्पर्धा}$ |
| $S. +,0$ | $IV. \text{असहभोजिता}$ |
| $T. +,-$ | $V. \text{सहभोजिता}$ |
| $VI. \text{परभक्षण}$ |
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